Strains of Angelfish
Most strains of angelfish available in the fishkeeping hobby are the result of many decades of selective breeding. For the most part, the original crosses of wild angelfish were not recorded and confusion between the various species of Pterophyllum, especially P. scalare and P. leopoldi, is common. This makes the origins of "Domestic angelfish" unclear. Domestic strains are most likely a collection of genes resulting from more than one species of wild angelfish combined with the selection of mutations in domesticated lines over the last 60 or more years. The result of this is a domestic angelfish that is a true hybrid with little more than a superficial resemblance to wild Pterophyllum species. It would be inaccurate to say that they accurately represent any species of wild angelfish, although they most resemble P. scalare and are frequently referred to as such.
Domestic angelfish have been bred and crossbred for several decades. There are hundreds of mutations of little importance by themselves. Much of the research into the known genetics of P. scalare is the result of the research of Dr. Joanne Norton, who published a series of 18 articles in Freshwater and Marine Aquarium (FAMA) Magazine. Those articles are reprinted at http://theangelfishsociety.org/genetics.htm .
Silver (+/+)
- The silver angelfish most commonly resembles the wild form of angelfish, and is also referred to as "wild-type". It is not, however, caught in the wild and is considered domestic. The fish has a silver body with red eyes and three vertical black stripes that can fade or darken depending on the mood of the fish.
Gold (g/g)
- The genetic trait for the gold angelfish is recessive, and causes a light golden body with a darker yellow or orange color on the crown of the fish. It does not have the vertical black stripes or the red eye seen in the wild angelfish.
Zebra (Z/+ or Z/Z)
- The zebra phenotype results in 4 to 6 vertical stripes on the fish that in other ways resembles a silver angelfish. It is a dominant mutation that exists at the same locus as the stripeless gene.
Black Lace (D/+) / Zebra Lace (D/+ - Z/+)
- A Silver or Zebra with one copy of the Dark gene. This results in very attractive lacing in the fins. Considered by some to the most attractive of all angelfish varieties.
Smokey (Sm/+)
- A variety with a dark brownish grey back half and dark dorsal and anal fins.
Chocolate (Sm/Sm)
- Homozygous for Smokey with more of the dark pattern. Sometimes only the head is silver.
Halfblack (h/h)
- Silver with a black rear portion. Halfblack can express along with some other color genes, but not all. The pattern may not develop or express if the fish are in stressful conditions.
Sunset Blushing (g/g S/S)
- The Sunset Blushing has two doses of gold and two doses of Stripeless. The upper half of the fish exhibits orange on the good ones. The body is mostly white in color, fins are clear. The amount of orange showing on the fish can vary. On some the body is a pinkish or tangerine color. The term blushing comes from the clear gill plates found on juveniles. You can see the pinkish gill underneath.
Koi (Gm/Gm S/S) or (Gm/g S/S)
- The Koi has a double or single dose of Gold Marble with a double dose of Stripeless. They express a variable amount of orange that varies with stress levels. The black marbling varies from 5%-40% coverage.
Leopard (Sm/Sm Z/Z) or (Sm/Sm Z/+)
- The leopard is a very popular fish when young, having spots over most of their body. Most of these spots grow closer together as an adult so it looks like a chocolate with dots on it. (Smokey x Zebra)
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